All of the flowers now raised here have developed from those _____ in the forest.    
A．once they grewB．they grew once
C．that once grewD．once grew
C
考查定語從句的關系詞的用法。分析句意和句子結構可知those之后是個定語從句，those相當于those flowers，是先行詞，因為是生長在森林里的（野）花，grew和關系代詞之間是主謂關系，因此that不能省。once在此表示“過去曾”，常與過去時連用，還可表示“每次”，引導從句時表示條件，翻譯成“一旦”。
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